Does Grice need an engram?

Dale excellently writes in that second 'historic' chapter to his dissertation on "The Theory of meaning", online:

"The gross differences between different causal theories are always about what sort of mental state gets caused by a public-language expression and what relation the adduced mental-state stands in to the meaning of the public-language expression. In Peirce's theory, a public-language expression causes what he called an interpretant<77>; in Russell's early theories, a mental image gets caused, as was just noted; for Ogden and Richards, it is what they call an engram."

--- as they drew from Greek!

I often wonder about Ogden and Richards.

Compare:

"Ogden's and Richards's use of 'engram'"

cfr.

"Grice's and Strawson's rejection of Quine's arguments".

I wonder if it was due to Richards -- or Ogden, the 'engram'.